The vertical control unit 70 includes a vertical combination control unit 71, a horizontal combination control unit 72 and an addition control unit 73. The vertical combination control unit 71 executes ON/OFF control of the first selection unit 16 (see FIG. 3) and the second selection unit 17 (see FIG. 3) in each pixel 10 so as to control combination processing executed to combine pixel signals from the plurality of pixels disposed consecutively along the column direction (vertical direction). The horizontal combination control unit 72 executes ON/OFF control of the plurality of switches constituting the horizontal combining unit 40 so as to control combination processing executed to combine pixel signals from the plurality of pixels disposed consecutively along the row direction (horizontal direction). The addition control unit 73 switches capacitors to be connected individually to the first vertical signal lines VL1 and the second vertical signal lines VL2 and to the arithmetic units 50 by controlling the ON/OFF states of a plurality of switches in the arithmetic units 50.
