The switch SW51 connects/disconnects the one terminal of the capacitor 51 with/from one terminal of the capacitor 53 and the switch SW52 connects/disconnects the one terminal of the capacitor 53 with/from one terminal of the capacitor 55. The switch SW53 connects/disconnects the one terminal of the capacitor 52 with/from the one terminal of the capacitor 54 and the switch SW54 connects/disconnects the one terminal of the capacitor 54 with/from one terminal of the capacitor 56. In addition, the switch SW55 connects/disconnects the one terminal of the capacitor 51 with/from the one terminal of the capacitor 55 and the switch SW56 connects/disconnects the one terminal of the capacitor 52 with/from the one terminal of the capacitor 56. Furthermore, the switch SW57 connects/disconnects the one terminal of the capacitor 51 with/from the one terminal of the capacitor 52 and the switch SW58 connects/disconnects the one terminal of the capacitor 53 with/from the one terminal of the capacitor 54. The switch SW59 connects/disconnects the one terminal of the capacitor 55 with/from the one terminal of the capacitor 56.
The arithmetic unit 50b outputs, through the output terminal 63, a signal generated by adding together signals input to the individual capacitors 51 through 56. The signal output from the arithmetic unit 50b first undergoes signal processing via an A/D conversion unit 80, the output unit 90 and the like and is then output to the control unit 4 of the camera 1. The following is a more detailed description of the pixel signal readout method adopted in the second control mode.