For example, the plurality of cover plate alignment marks comprise four cover plate alignment marks, the four cover plate alignment marks are respectively numbered as a first cover plate alignment mark, a second cover plate alignment mark, a third cover plate alignment mark, and a fourth cover plate alignment mark in an anticlockwise direction; with respect to an axis parallel to the first direction, a combination of the first cover plate alignment mark and the second cover plate alignment mark is symmetrical with a combination of the third cover plate alignment mark and the fourth cover plate alignment mark.
For example, the display substrate further comprises an electrode layer extending portion electrically connected with the plurality of light emitting elements, and the electrode layer extending portion is in the frame region; in the direction perpendicular to the base substrate, the electrode layer extending portion is between the plurality of cover plate alignment marks and the base substrate; and an orthographic projection of the electrode layer extending portion on the base substrate is overlapped with orthographic projections of the plurality of cover plate alignment marks on the base substrate.
For example, the electrode layer extending portion comprises a first conductive layer and a second conductive layer which are stacked, the second conductive layer is on a side of the first conductive layer away from the base substrate, and the second conductive layer comprises a portion recessed toward the base substrate, the portion is electrically connected with the first conductive layer, and the plurality of cover plate alignment marks are in a recessed region defined by the portion.